Cole, C.J.. „Amphibians and reptiles of Guyana, South America: Illustrated keys, annotated species accounts, and a biogeographic synopsis”. Proceedings of the Biological Society of Washington125, 317–620. o. DOI:10.2988/0006-324x-125.4.317.
Fouquet (2012. február 20.). „From Amazonia to the Atlantic forest: Molecular phylogeny of Phyzelaphryninae frogs reveals unexpected diversity and a striking biogeographic pattern emphasizing conservation challenges”. Molecular Phylogenetics and Evolution65, 547–561. o. DOI:10.1016/j.ympev.2012.07.012. PMID22842094.
Fouquet (2012. február 20.). „From Amazonia to the Atlantic forest: Molecular phylogeny of Phyzelaphryninae frogs reveals unexpected diversity and a striking biogeographic pattern emphasizing conservation challenges”. Molecular Phylogenetics and Evolution65, 547–561. o. DOI:10.1016/j.ympev.2012.07.012. PMID22842094.