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Wellman, Charles H. (2018), "Palaeoecology and palaeophytogeography of the Rhynie chert plants: Further evidence from integrated analysis of in situ and dispersed spores", Philosophical Transactions of the Royal Society B (dalam bahasa Inggris), 373 (1739): 20160491, doi:10.1098/rstb.2016.0491, PMC5745327, PMID29254956