Carnevale, Giorgio; Pietsch, Theodore W. (2010). "Eocene handfishes from Monte Bolca, with description of a new genus and species, and a phylogeny of the family Brachionichthyidae (Teleostei: Lophiiformes)". Zoological Journal of the Linnean Society. 160 (4): 621–647. doi:10.1111/j.1096-3642.2009.00623.x.