Lai, JCY., PKL. Ng & PJF. Davie. "A revision of the Portunus pelagicus (Linnaeus, 1758) species complex (Crustacea: Brachyura: Portunidae), with the recognition of four species." The Raffles Bulletin of Zoology58(2): 199-237.Diarsipkan 2014-04-22 di Wayback Machine. (31 Agustus 2010)
Lai, JCY., PKL. Ng & PJF. Davie. "A revision of the Portunus pelagicus (Linnaeus, 1758) species complex (Crustacea: Brachyura: Portunidae), with the recognition of four species." The Raffles Bulletin of Zoology58(2): 199-237.Diarsipkan 2014-04-22 di Wayback Machine. (31 Agustus 2010)