Sasaki T, Kartavtsev YP, Chiba SN, Uematsu T, Sviridov VV, Hanzawa N (2007). “Genetic divergence and phylogenetic independence of Far Eastern species in subfamily Leuciscinae (Pisces: Cyprinidae) inferred from mitochondrial DNA analyses”. Genes Genet. Syst.82 (4): 329-340. doi:10.1266/ggs.82.329.
Silvia Perea, Madelaine Böhme, Primož Zupančič, Jörg Freyhof, Radek Šanda, Müfit Özuluğ, Asghar Abdoli,Ignacio Doadrio (2010). “Phylogenetic relationships and biogeographical patterns in Circum-Mediterranean subfamily Leuciscinae (Teleostei, Cyprinidae) inferred from both mitochondrial and nuclear data”. BMC Evolutionary Biology10. doi:10.1186/1471-2148-10-265.
Rafael Zardoya and Ignacio Doadrio (1998). “Phylogenetic relationships of Iberian cyprinids:systematic and biogeographical implications”. Proc. R. Soc. Lond. B265 (1403): 1365-1372. doi:10.1098/rspb.1998.0443.
“Phylogenetic relationships of Eurasian and American cyprinids using cytochrome b sequences”. Journal of Fish Biology61: 929–944. (2002). doi:10.1111/j.1095-8649.2002.tb01853.x.
Derek D. Houston, Dennis K. Shiozawa, Brett R. Riddle (2010). “Phylogenetic relationships of the western North American cyprinid genus Richardsonius, with an overview of phylogeographic structure”. Mol. Phyl. Evol.55: 259-273. doi:10.1016/j.ympev.2009.10.017.
ANDREW M. SIMONS, PETER B. BERENDZEN and RICHARD L. MAYDEN (2003). “Molecular systematics of North American phoxinin genera (Actinopterygii: Cyprinidae) inferred from mitochondrial 12S and 16S ribosomal RNA sequences”. Zool J Linn Soc.139: 63-80. doi:10.1046/j.1096-3642.2003.00076.x.
fishbase.org
Froese, Rainer, and Daniel Pauly, eds. (2006). "Cyprinidae" in FishBase. April 2006 version.