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Stelbrink, B.; von Rintelen, T.; Cliff, G.; Kriwet, J. (2010). “Molecular systematics and global phylogeography of angel sharks (genus Squatina)”. Molecular Phylogenetics and Evolution54 (2): 395–404. doi:10.1016/j.ympev.2009.07.029. PMID19647086.
Nagasawa, K.; Tanaka, S.; Benz, G.W. (1998). “Trebius shiinoi n. sp. (Trebiidae: Siphonostomatoida: Copepoda) from uteri and embryos of the Japanese angelshark (Squatina japonica) and the clouded angelshark (Squatina nebulosa), and redescription of Trebius longicaudatus”. Journal of Parasitology84 (6): 1218–1230. doi:10.2307/3284678. PMID9920318.