Klaus-Peter Koepfli, Susan M. Jenks, Eduardo Eizirik, Tannaz Zahirpour, Blaire Van Valkenburgh, Robert K. Wayne (2006). “Molecular systematics of the Hyaenidae: Relationships of a relictual lineage resolved by a molecular supermatrix”. Molecular Phylogenetics and Evolution38 (3): 603-620. doi:10.1016/j.ympev.2005.10.017.
Turner, Alan; Antón, Mauricio; Werdelin, Lars (2008). “Taxonomy and evolutionary patterns in the fossil Hyaenidae of Europe”. Geobios41 (5): 677–687. doi:10.1016/j.geobios.2008.01.001.
Lars Werdelin & Nikos Solounias, “The Hyaenidae: taxonomy, systematics and evolution,” Fossils and Strata, No. 30, Scandinavian University Press, 1991, Pages 1–104. https://doi.org/10.18261/8200374815-1991-01.
