Hu, Tao & Corey, E.J. (2002). “Short Syntheses of (±)-δ-Araneosene and Humulene Utilizing a Combination of Four-Component Assembly and Palladium-Mediated Cyclization”. Organic Letters4 (14): 2441-2443. doi:10.1021/ol026205p. PMID12098267.
Yange, Xiaogen; Lederer, Cindy; McDaniel, Mina & Deinzer, Max. (1993). “Evaluation of hydrolysis products of humulene epoxides II and III”. Journal of Agricultural and Food Chemistry41 (8): 1300-1304. doi:10.1021/jf00032a026.
Fernandes E.S., Passos G.F., Medeiros R., da Cunha F.M., Ferreira J., Campos M.M., Pianowski L.F., Calixto J.B. (2007). “Anti-inflammatory effects of compounds alpha-humulene and (-)-trans-caryophyllene isolated from the essential oil of Cordia verbenacea”. European Journal of Pharmacology569 (3): 228-236. doi:10.1016/j.ejphar.2007.04.059. PMID17559833.
enzyme-database.org
Moss, G.P., “Humulene derived sesquiterpenoid biosynthesis.” International Union of Biochemistry and Molecular Biology Enzyme Nomenclature. Accessed April 10, 2011. http://www.enzyme-database.org/reaction/terp/humul.html
Hu, Tao & Corey, E.J. (2002). “Short Syntheses of (±)-δ-Araneosene and Humulene Utilizing a Combination of Four-Component Assembly and Palladium-Mediated Cyclization”. Organic Letters4 (14): 2441-2443. doi:10.1021/ol026205p. PMID12098267.
Fernandes E.S., Passos G.F., Medeiros R., da Cunha F.M., Ferreira J., Campos M.M., Pianowski L.F., Calixto J.B. (2007). “Anti-inflammatory effects of compounds alpha-humulene and (-)-trans-caryophyllene isolated from the essential oil of Cordia verbenacea”. European Journal of Pharmacology569 (3): 228-236. doi:10.1016/j.ejphar.2007.04.059. PMID17559833.
realbeer.com
Glenn Tinseth, "Hop Aroma and Flavor", January/February 1993, Brewing Techniques. <http://realbeer.com/hops/aroma.html> Accessed July 21, 2010