De Carvalho, M.R.; M.P. Ragno (2011). “An unusual, dwarf species of Neotropical freshwater stingray, Plesiotrygon nana sp. nov., from the upper and mid Amazon basin: the second species of Plesiotrygon (Chondrichthyes: Potamotrygonidae)”. Papéis Avulsos de Zoologia51 (7): 101–138. doi:10.1590/S0031-10492011000700001.