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Ritland, D.; L. P. Brower (1991). “The viceroy butterfly is not a Batesian mimic”. Nature350 (6318): 497–498. Bibcode: 1991Natur.350..497R. doi:10.1038/350497a0. "Viceroys are as unpalatable as monarchs, and significantly more unpalatable than queens from representative Florida populations."
Wilson, Joshua S.; Williams, Kevin A.; Forister, Matthew L.; von Dohlen, Carol D.; Pitts, James P. (2012). “Repeated evolution in overlapping mimicry rings among north American velvet ants”. Nature Communications3: 1272. Bibcode: 2012NatCo...3.1272W. doi:10.1038/ncomms2275. PMID23232402.
Mullen, S.P. (2006). “Wing pattern evolution and the origins of mimicry among North American admiral butterflies (Nymphalidae: Limenitis)”. Molecular Phylogenetics and Evolution39 (3): 747–758. doi:10.1016/j.ympev.2006.01.021. PMID16500119.
Wilson, Joshua S.; Williams, Kevin A.; Forister, Matthew L.; von Dohlen, Carol D.; Pitts, James P. (2012). “Repeated evolution in overlapping mimicry rings among north American velvet ants”. Nature Communications3: 1272. Bibcode: 2012NatCo...3.1272W. doi:10.1038/ncomms2275. PMID23232402.