Titov, V. V.; Shvyreva, A. K. (January 2016). “Deer of the genus Megaloceros (Mammalia, Cervidae) from the Early Pleistocene of Ciscaucasia” (英語). Paleontological Journal50 (1): 87–95. doi:10.1134/S0031030116010111. ISSN0031-0301.
Titov, V. V.; Shvyreva, A. K. (January 2016). “Deer of the genus Megaloceros (Mammalia, Cervidae) from the Early Pleistocene of Ciscaucasia” (英語). Paleontological Journal50 (1): 87–95. doi:10.1134/S0031030116010111. ISSN0031-0301.