Lister, Adrian M.; Edwards, Ceiridwen J.; Nock, D. A. W.; Bunce, Michael; van Pijlen, Iris A.; Bradley, Daniel G.; Thomas, Mark G.; Barnes, Ian (2005). “The phylogenetic position of the giant deer Megaloceros giganteus”. ネイチャー438 (7069): 850–853. Bibcode: 2005Natur.438..850L. doi:10.1038/nature04134. PMID16148942.
Gonzalez, Silvia; Kitchener, Andrew C.; Lister, Adrian (2000-07). “Survival of the Irish Elk into the Holocene”. ネイチャー405 (6788): 753-754. doi:10.1038/35015668.
Van der Made, Jan (2019-06-20). “The dwarfed "giant deer" Megaloceros matritensis n.sp. from the Middle Pleistocene of Madrid - A descendant of M. savini and contemporary to M. giganteus”. Quaternary International(英語版)520: 110-139. doi:10.1016/j.quaint.2018.06.006.
Lister, Adrian M.; Edwards, Ceiridwen J.; Nock, D. A. W.; Bunce, Michael; van Pijlen, Iris A.; Bradley, Daniel G.; Thomas, Mark G.; Barnes, Ian (2005). “The phylogenetic position of the giant deer Megaloceros giganteus”. ネイチャー438 (7069): 850–853. Bibcode: 2005Natur.438..850L. doi:10.1038/nature04134. PMID16148942.
Lister, Adrian M.; Edwards, Ceiridwen J.; Nock, D. A. W.; Bunce, Michael; van Pijlen, Iris A.; Bradley, Daniel G.; Thomas, Mark G.; Barnes, Ian (2005). “The phylogenetic position of the giant deer Megaloceros giganteus”. ネイチャー438 (7069): 850–853. Bibcode: 2005Natur.438..850L. doi:10.1038/nature04134. PMID16148942.
Van der Made, Jan (2019-06-20). “The dwarfed "giant deer" Megaloceros matritensis n.sp. from the Middle Pleistocene of Madrid - A descendant of M. savini and contemporary to M. giganteus”. Quaternary International(英語版)520: 110-139. doi:10.1016/j.quaint.2018.06.006.