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Hay, Jennifer M.; Sarre, Stephen D.; Lambert, David M.; Allendorf, Fred W.; Daugherty, Charles H. (June 2010). “Genetic diversity and taxonomy: a reassessment of species designation in tuatara (Sphenodon: Reptilia)”. Conservation Genetics11 (3): 1063–1081. doi:10.1007/s10592-009-9952-7. hdl:10072/30480.