Koepfli, K.-P.; Jenks, S. M.; Eizirik, E.; Zahirpour, T.; Van Valkenburgh, B.; Wayne, R. K. (2006). “Molecular systematics of the Hyaenidae: Relationships of a Relictual Lineage Resolved by a Molecular Supermatrix”. 《Molecular Phylogenetics and Evolution》 38 (3): 603–620. Bibcode:2006MolPE..38..603K. CiteSeerX10.1.1.529.1977. doi:10.1016/j.ympev.2005.10.017. PMID16503281.
Klaus-Peter Koepfli, Susan M. Jenks, Eduardo Eizirik, Tannaz Zahirpour, Blaire Van Valkenburgh, Robert K. Wayne: Molecular systematics of the Hyaenidae: Relationships of a relictual lineage resolved by a molecular supermatrix. In: Molecular Phylogenetics and Evolution. 38, 2006, S. 603–620 und Anne D. Yoder, Melissa M. Burns, Sarah Zehr, Thomas Delefosse, Geraldine Veron, Steven M. Goodman, John J. Flynn: Single origin of Malagasy Carnivora from an African ancestor. (PDF; 388 kB)
Koepfli, K.-P.; Jenks, S. M.; Eizirik, E.; Zahirpour, T.; Van Valkenburgh, B.; Wayne, R. K. (2006). “Molecular systematics of the Hyaenidae: Relationships of a Relictual Lineage Resolved by a Molecular Supermatrix”. 《Molecular Phylogenetics and Evolution》 38 (3): 603–620. Bibcode:2006MolPE..38..603K. CiteSeerX10.1.1.529.1977. doi:10.1016/j.ympev.2005.10.017. PMID16503281.
Koepfli, K.-P.; Jenks, S. M.; Eizirik, E.; Zahirpour, T.; Van Valkenburgh, B.; Wayne, R. K. (2006). “Molecular systematics of the Hyaenidae: Relationships of a Relictual Lineage Resolved by a Molecular Supermatrix”. 《Molecular Phylogenetics and Evolution》 38 (3): 603–620. Bibcode:2006MolPE..38..603K. CiteSeerX10.1.1.529.1977. doi:10.1016/j.ympev.2005.10.017. PMID16503281.
Koepfli, K.-P.; Jenks, S. M.; Eizirik, E.; Zahirpour, T.; Van Valkenburgh, B.; Wayne, R. K. (2006). “Molecular systematics of the Hyaenidae: Relationships of a Relictual Lineage Resolved by a Molecular Supermatrix”. 《Molecular Phylogenetics and Evolution》 38 (3): 603–620. Bibcode:2006MolPE..38..603K. CiteSeerX10.1.1.529.1977. doi:10.1016/j.ympev.2005.10.017. PMID16503281.