Roskov Y., Kunze T., Orrell T., Abucay L., Paglinawan L., Culham A., Bailly N., Kirk P., Bourgoin T., Baillargeon G., Decock W., De Wever A., Didžiulis V. (ed) (2014). “Species 2000 & ITIS Catalogue of Life: 2014 Annual Checklist.”. Species 2000: Reading, UK. 2015년 8월 6일에 확인함. CS1 관리 - 여러 이름 (링크)
doi.org
dx.doi.org
Villarreal; 외. (2016). “Divergence times and the evolution of morphological complexity in an early land plant lineage (Marchantiopsida) with a slow molecular rate”. 《New Phytologist》 209 (4): 1734–46. doi:10.1111/nph.13716. PMID26505145.
nih.gov
ncbi.nlm.nih.gov
Villarreal; 외. (2016). “Divergence times and the evolution of morphological complexity in an early land plant lineage (Marchantiopsida) with a slow molecular rate”. 《New Phytologist》 209 (4): 1734–46. doi:10.1111/nph.13716. PMID26505145.