Kuroiwa A, Handa S, Nishiyama C, Chiba E, Yamada F, Abe S, Matsuda Y (2011년 6월 8일). “Additional copies of CBX2 in the genomes of males of mammals lacking SRY, the Amami spiny rat (Tokudaia osimensis) and the Tokunoshima spiny rat (Tokudaia tokunoshimensis)”. 《Chromosome Res》 19: 635–44. doi:10.1007/s10577-011-9223-6. PMID21656076.
Kuroiwa A, Handa S, Nishiyama C, Chiba E, Yamada F, Abe S, Matsuda Y (2011년 6월 8일). “Additional copies of CBX2 in the genomes of males of mammals lacking SRY, the Amami spiny rat (Tokudaia osimensis) and the Tokunoshima spiny rat (Tokudaia tokunoshimensis)”. 《Chromosome Res》 19: 635–44. doi:10.1007/s10577-011-9223-6. PMID21656076.