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Eizirik, E.; Murphy, W.J.; Koepfli, K.P.; Johnson, W.E.; Dragoo, J.W.; O'Brien, S.J. (2010). “Pattern and timing of the diversification of the mammalian order Carnivora inferred from multiple nuclear gene sequences”. 《Molecular Phylogenetics and Evolution》 56: 49–63. doi:10.1016/j.ympev.2010.01.033. PMID20138220.
