Roehrs, Z.P.; Lack, J.B.; Van Den Bussche, R.A. (2010). “Tribal phylogenetic relationships within Vespertilioninae (Chiroptera: Vespertilionidae) based on mitochondrial and nuclear sequence data”. 《Journal of Mammalogy》 91 (5): 1073–1092. doi:10.1644/09-MAMM-A-325.1.