Chomicki, Guillaume; Bidel, Luc P. R.; Ming, Feng; 외. (2015). “The velamen protects photosynthetic orchid roots against UV-B damage, and a large dated phylogeny implies multiple gains and losses of this function during the Cenozoic”. 《New Phytologist》 (영어) 205 (3): 1330–1341. doi:10.1111/nph.13106.
Givnish, Thomas J.; Spalink, Daniel; Ames, Mercedes; 외. (2015). “Orchid phylogenomics and multiple drivers of their extraordinary diversification”. 《Proceedings of the Royal Society B: Biological Sciences》 (영어) 282 (1814). doi:10.1098/rspb.2015.1553.
