Rull, Valenti (2005년 5월 18일). “Biotic diversification in the Guayana Highlands: a proposal”. 《Journal of Biogeography》 32 (6): 921–927. doi:10.1111/j.1365-2699.2005.01252.x.
Salerno, P. E.; Ron, S. R.; Señaris, J. C.; Rojas-Runjaic, F. J. M.; Noonan, B. P.; Cannatella, D. C. (2012). “Ancient Tepui Summits Harbor Young Rather Than Old Lineages of Endemic Frogs”. 《Evolution》 66 (10): 3000–3013. doi:10.1111/j.1558-5646.2012.01666.x. PMID23025594. S2CID205783918.
nih.gov
ncbi.nlm.nih.gov
Salerno, P. E.; Ron, S. R.; Señaris, J. C.; Rojas-Runjaic, F. J. M.; Noonan, B. P.; Cannatella, D. C. (2012). “Ancient Tepui Summits Harbor Young Rather Than Old Lineages of Endemic Frogs”. 《Evolution》 66 (10): 3000–3013. doi:10.1111/j.1558-5646.2012.01666.x. PMID23025594. S2CID205783918.
Salerno, P. E.; Ron, S. R.; Señaris, J. C.; Rojas-Runjaic, F. J. M.; Noonan, B. P.; Cannatella, D. C. (2012). “Ancient Tepui Summits Harbor Young Rather Than Old Lineages of Endemic Frogs”. 《Evolution》 66 (10): 3000–3013. doi:10.1111/j.1558-5646.2012.01666.x. PMID23025594. S2CID205783918.