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R. M. Bush; W. M. Fitch; C. A. Bender; N. J. Cox (1999). “Positive selection on the H3 hemagglutinin gene of human influenza virus”. 《Molecular Biology and Evolution》 16 (11): 1457–1465. doi:10.1093/oxfordjournals.molbev.a026057. PMID10555276.