Giannini, Norberto P.; Almeida, Francisca Cunha; Simmons, Nancy B.; Helgen, Kristofer M. (2008). “The systematic position of Pteropus leucopterus and its bearing on the monophyly and relationships of Pteropus (Chiroptera : Pteropodidae)”. 《Acta Chiropterologica》 10: 11–20. doi:10.3161/150811008x331054.