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McLellan, B.N., Proctor, M.F., Huber, D. & Michel, S. 2017. Ursus arctos (amended version of 2017 assessment). The IUCN Red List of Threatened Species 2017: e.T41688A121229971. https://dx.doi.org/10.2305/IUCN.UK.2017-3.RLTS.T41688A121229971.en. Downloaded on 27 April 2021.
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Calvignac, S.; Hughes, S.; Tougard, C.; Michaux, J.; Thevenot, M.; Philippe, M.; Hamdine, W.; Hanni, C. (2008). "Ancient DNA evidence for the loss of a highly divergent brown bear clade during historical times". Mol. Ecol. 17 (8): 1962–1970. doi:10.1111/j.1365-294x.2008.03631.x. PMID18363668.