Marin, M.A., Peña, C., Freitas, A.V.L., Wahlberg, N. & Uribe, S.I. (2011). From the phylogeny of the Satyrinae butterflies to the systematics of Euptychiina (Lepidoptera, Nymphalidae): history, progress and prospects. Neotropical Entomology40: 1-13.Gearchiveerd op 10 januari 2010.
web.archive.org
Peña, C., N. Wahlberg, E. Weingartner, U. Kodandaramaiah, S. Nylin, A. V. L. Freitas, & A. V. Z. Brower (2006). "Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data". Molecular Phylogenetics and Evolution40: 29-49.
Marin, M.A., Peña, C., Freitas, A.V.L., Wahlberg, N. & Uribe, S.I. (2011). From the phylogeny of the Satyrinae butterflies to the systematics of Euptychiina (Lepidoptera, Nymphalidae): history, progress and prospects. Neotropical Entomology40: 1-13.Gearchiveerd op 10 januari 2010.
