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(en) William T. White, Shannon Corrigan, Lei Yang, Aaron C. Henderson, Adam L. Bazinet, David L. Swofford, Gavin J. P. Naylor(January 2018).Phylogeny of the manta and devilrays (Chondrichthyes: mobulidae), with an updated taxonomic arrangement for the family. Zoological Journal of the Linnean Society182(1): 50–75. DOI: 10.1093/zoolinnean/zlx018.
