(en) Ronquist F, Klopfstein S, Vilhelmsen L. (2012). A Total-Evidence Approach to Dating with Fossils, Applied to the Early Radiation of the Hymenoptera. Systematic Biology61 (6): 973–999. PMID22723471. DOI: 10.1093/sysbio/sys058.
(en) Janke A, Klopfstein S, Vilhelmsen L. et al. (2013). The Hymenopteran Tree of Life: Evidence from Protein-Coding Genes and Objectively Aligned Ribosomal Data. PLoS ONE8 (8): e69344. PMID23936325. PMC3732274. DOI: 10.1371/journal.pone.0069344.
(en) Ronquist F, Klopfstein S, Vilhelmsen L. (2012). A Total-Evidence Approach to Dating with Fossils, Applied to the Early Radiation of the Hymenoptera. Systematic Biology61 (6): 973–999. PMID22723471. DOI: 10.1093/sysbio/sys058.
(en) Janke A, Klopfstein S, Vilhelmsen L. et al. (2013). The Hymenopteran Tree of Life: Evidence from Protein-Coding Genes and Objectively Aligned Ribosomal Data. PLoS ONE8 (8): e69344. PMID23936325. PMC3732274. DOI: 10.1371/journal.pone.0069344.