Woo, Patrick C. Y.; Huang, Yi; Lau, Susanna K. P.; Yuen, Kwok-Yung (24. august 2010). «Coronavirus Genomics and Bioinformatics Analysis». Viruses. 2 (8): 1804–1820. PMC3185738. PMID21994708. doi:10.3390/v2081803. «In all members of Betacoronavirus subgroup A, a haemagglutinin esterase (HE) gene, which encodes a glycoprotein with neuraminate O-acetyl-esterase activity and the active site FGDS, is present downstream to ORF1ab and upstream to S gene (Figure 1).»
Woo, Patrick C. Y.; Huang, Yi; Lau, Susanna K. P.; Yuen, Kwok-Yung (24. august 2010). «Coronavirus Genomics and Bioinformatics Analysis». Viruses. 2 (8): 1804–1820. PMC3185738. PMID21994708. doi:10.3390/v2081803. «In all members of Betacoronavirus subgroup A, a haemagglutinin esterase (HE) gene, which encodes a glycoprotein with neuraminate O-acetyl-esterase activity and the active site FGDS, is present downstream to ORF1ab and upstream to S gene (Figure 1).»