David A. S. Smith, Gugs Lushai, John A. Allen. A classification of Danaus butterflies (Lepidoptera: Nymphalidae) based upon data from morphology and DNA. „Zoological Journal of the Linnean Society”. 144 (2), s. 191–212, 2005. DOI: 10.1111/j.1096-3642.2005.00169.x.
Traut, Walther & Ahola, Virpi & A.S. Smith, David & Gordon, Ian & ffrench-Constant, Richard. Karyotypes versus Genomes: The Nymphalid Butterflies Melitaea cinxia, Danaus plexippus, and D. chrysippus.. „Cytogenetic and Genome Research”. 153, 2017. DOI: 10.1159/000484032.
Genus Danaus, Kluk 1780. W: Mark C. Williams: Afrotropical Butterflies. Wyd. 17. Lepidopterists Society of Africa, 2018. (ang.). Brak numerów stron w książce