Darrel R.D.R.FrostDarrel R.D.R., Holoadeninae Hedges, Duellman, and Heinicke, 2008, [w:] Amphibian Species of the World: an Online Reference. Version 6.2 [online], American Museum of Natural History, New York, USA [dostęp 2025-06-03](ang.).
doi.org
M.P. Heinicke, A.R. Lemmon, E.M. Lemmon, K. McGrath & S.B. Hedges. Phylogenomic support for evolutionary relationships of New World direct-developing frogs (Anura: Terraranae). „Molecular Phylogenetics and Evolution”. 118, s. 145, 2018. DOI: 10.1016/j.ympev.2017.09.021. (ang.).