Andrzej Szeptycki. The taxonomy of Protura – present status and future problems. „Pedobiologia”. 46 (3-4), s. 209–214, 2002. DOI: 10.1078/0031-4056-00127. (ang.).
Luan Yuanxia, John M. Mallatt, Xie Rongdong, Yang Yiming, Yin Wenying. The phylogenetic positions of three basal-hexapod groups (Protura, Diplura, and Collembola) based on ribosomal RNA gene sequences. „Molecular Biology and Evolution”. 22 (7), s. 1579–1592, 2005. DOI: 10.1093/molbev/msi148. (ang.).