D.E. Wilson & D.M. Reeder (redaktorzy): Genus Spermophilus. [w:] Mammal Species of the World. A Taxonomic and Geographic Reference (Wyd. 3) [on-line]. Johns Hopkins University Press, 2005. [dostęp 2020-12-29].
doi.org
doi.org
E. Simonov, N.V.Lopatina, S.V. Titov, A.D. Ivanova, O.V. Brandler, V.L. Surin, V.A. Matrosova, A.E. Dvilis, N.V. Oreshkova, S.Y. Kapustina, F.N. Golenishchev & O.A. Ermakov. Traditional multilocus phylogeny fails to fully resolve Palearctic ground squirrels (Spermophilus) relationships but reveals a new species endemic to West Siberia. „ПMolecular Phylogenetics and Evolution”. 195, s. 108057, 2024. DOI: 10.1016/j.ympev.2024.108057. (ang.).
Maxim V Sinitsa, Natalia V. Pogodina, Lyudmila Y. Кryuchkova. The skull of Spermophilus nogaici (Rodentia: Sciuridae: Xerinae) and the affinities of the earliest Old World ground squirrels. „Zoological Journal of the Linnean Society”. 186 (3), s. 826–864, 2019. DOI: 10.1093/zoolinnean/zly092. (ang.).
M.V. Sinitsa & N.V. Pogodina. The evolution of early Spermophilus in eastern Europe and the antiquity of the Old World ground squirrels. „Acta Palaeontologica Polonica”. 64 (3), s. 643–667, 2019. DOI: 10.4202/app.00605.2019. (ang.).
N. Upham, C. Burgin, J. Widness, M. Becker, J. Zijlstra & D. Huckaby: Treeview of Mammalian Taxonomy Hierarchy. [w:] ASM Mammal Diversity Database (Version 1.13) [on-line]. American Society of Mammalogists. [dostęp 2024-07-18]. (ang.).