Artiss et al. Molecular phylogenetic analysis of the dragonfly genera Libellula, Ladona, and Plathemis (Odonata: Libellulidae) based on mitochondrial cytochrome oxidase I and 16S rRNA sequence data. „Molecular Phylogenetics and Evolution”. 18 (3), s. 348–361, 2001. DOI: 10.1006/mpev.2000.0867. (ang.).