Hirata R, Ohsumk Y, Nakano A, Kawasaki H, Suzuki K, Anraku Y (abril de 1990). «Molecular structure of a gene, VMA1, encoding the catalytic subunit of H(+)-translocating adenosine triphosphatase from vacuolar membranes of Saccharomyces cerevisiae». J. Biol. Chem. 265 (12): 6726–33. PMID2139027