Luis A. Coloma, Santiago Ron, Martín R. Bustamante, Diego Cisneros-Heredia, Ana Almendáriz 2004. Atelopus guanujo. In: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.1. Downloaded on 14 September 2012.
apiv3.iucnredlist.org
Luis A. Coloma, Santiago Ron, Martín R. Bustamante, Diego Cisneros-Heredia, Ana Almendáriz 2004. Atelopus guanujo. In: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.1. Downloaded on 14 September 2012.