Alexander W. A. Kellner, Xiaolin Wang, Helmut Tischlinger, Diogenes de Almeida Campos, David W. E. Hone, and Xi Meng. (2009). "The soft tissue of Jeholopterus (Pterosauria, Anurognathidae, Batrachognathinae) and the structure of the pterosaur wing membrane", Proc. R. Soc. Bdoi:10.1098/rspb.2009.0846