Arden, T. M. S.; Klein, C. G.; Zouhri, S.; Longrich, N. R. (2019). «Aquatic adaptation in the skull of carnivorous dinosaurs (Theropoda: Spinosauridae) and the evolution of aquatic habits in Spinosaurus». Cretaceous Research. 93: 275–284. doi:10.1016/j.cretres.2018.06.013