Carvalho, A.L.G. (2016). «Three new species of the Tropidurus spinulosus group (Squamata: Tropiduridae) from eastern Paraguay.». American Museum Novitates. 3853: 1-44. doi:10.1206/3853.1
de Carvalho, A.L.G., de Britto M.R. and Fernandes D.S. (2013). «Biogeography of the lizard genus Tropidurus Wied- Neuwied, 1825 (Squamata: Tropiduridae): distribution, endemism, and area relationships in South America». PLoS ONE. 8 (3): e59736. doi:10.1371/journal.pone.0059736 !CS1 manut: Nomes múltiplos: lista de autores (link)
Cacciali, P., Scott, N.J., Luz Aquino Ortíz, A. Fitzgerald, L., Smith, P. (2016). «The Reptiles of Paraguay: Literature, Distribution, and an Annotated Taxonomic Checklist». Museum of Southwestern Biology. ISSN2380-4599 !CS1 manut: Nomes múltiplos: lista de autores (link)