Lovegrove, B. G. (). „Cool sperm: Why some placental mammals have a scrotum”. Journal of Evolutionary Biology. 27 (5): 801–814. doi:10.1111/jeb.12373. PMID24735476.
Liguori, Giovanni; Ollandini, Giangiacomo; Napoli, Renata; Mazzon, Giorgio; Petrovic, Miloš; Trombetta, Carlo (), Bertolotto, Michele; Trombetta, Carlo, ed., „Anatomy of the Scrotum”, Scrotal Pathology, Springer Berlin Heidelberg, pp. 27–34, doi:10.1007/174_2011_170, ISBN978-3-642-12455-6, accesat în
Gallup, Gordon G.; Finn, Mary M.; Sammis, Becky (). „On the Origin of Descended Scrotal Testicles: The Activation Hypothesis”. Evolutionary Psychology. 7 (4): 147470490900700. doi:10.1177/147470490900700402.
Lovegrove, B. G. (). „Cool sperm: Why some placental mammals have a scrotum”. Journal of Evolutionary Biology. 27 (5): 801–814. doi:10.1111/jeb.12373. PMID24735476.