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Wahlberg N., Braby M.F., Brower A.V., et al. Synergistic effects of combining morphological and molecular data in resolving the phylogeny of butterflies and skippers (англ.) // Proceedings of the Royal Society B : journal. — 2005. — August (vol. 272, no. 1572). — P. 1577—1586. — doi:10.1098/rspb.2005.3124. — PMID16048773. — PMC1560179.