Zinenko O., Stümpel N., Mazanaeva L., Bakiev A., Shiryaev K., Pavlov A., Kotenko T., Kukushkin O., Chikin Yu., Duisebajeva T., Nilson G., Orlov N. L., Tuniyev S., Ananjeva N. B., Murphy R. W., Joger U. Mitochondrial phylogeny shows multiply independent ecological transitions and northern dispersion despite of Pleistocene glaciations in meadow and steppe vipers (Vipera ursinii and Vipera renardi) : [англ.] // Molecular Phylogenetics and Evolution. — 2015. — Vol. 84. — P. 85—100. — doi:10.1016/j.ympev.2014.12.005.
Freitas I., Ursenbacher S., Mebert K., Zinenko O., Schweiger S., Wüster W., Brito J., Crnobrnja-Isailovic J., Halpern B., Fahd S., Santos X., Pleguezuelos J. M., Joger U., Orlov N., Mizsei E., Lourdais O., Zuffi M. A. L., Strugariu A., Zamfirescu ¸S. R., Martínez-Solano Í., Velo-Antón G., Kaliontzopoulou A., Martínez-Freiría F. Evaluating taxonomic inflation: towards evidence-based species delimitation in Eurasian vipers (Serpentes: Viperinae) : [англ.] // Amphibia-Reptilia. — 2020. — Vol. 41, no. 3. — P. 285–311. — doi:10.1163/15685381-bja10007.
