Yongshuai Sun, Richard J. Abbott, Lili Li, Long Li, Jiabin Zou & Jianquan Liu. Evolutionary history of Purple cone spruce (Picea purpurea) in the Qinghai–Tibet Plateau: homoploid hybrid origin and Pleistocene expansion (англ.) // Molecular Ecology[англ.] : journal. — 2014. — Vol. 23, no. 2. — P. 343—359. — doi:10.1111/mec.12599.
Yongshuai Sun, Richard J. Abbott, Lili Li, Long Li, Jiabin Zou & Jianquan Liu. Evolutionary history of Purple cone spruce (Picea purpurea) in the Qinghai–Tibet Plateau: homoploid hybrid origin and Pleistocene expansion (англ.) // Molecular Ecology[англ.] : journal. — 2014. — Vol. 23, no. 2. — P. 343—359. — doi:10.1111/mec.12599.