Artiss T., Schultz T.R., Polhemus D.A., Simon C. Molecular phylogenetic analysis of the dragonfly genera Libellula, Ladona, and Plathemis (Odonata: Libellulidae) based on mitochondrial cytochrome oxidase I and 16S rRNA sequence data (англ.) // Molecular Phylogenetics and Evolution : journal. — Academic Press, 2001. — Vol. 18, no. 3. — P. 348—361. — doi:10.1006/mpev.2000.0867. — PMID11277629.
Kambhampati, Srinivas; Charlton, Ralph E. Phylogenetic relationship among Libellula, Ladona and Plathemis (Odonata: Libellulidae) based on DNA sequence of mitochondrial 16S rRNA gene (англ.) // Systematic Entomology : journal. — Wiley-Blackwell, 1999. — Vol. 24, no. 1. — P. 37—49. — doi:10.1046/j.1365-3113.1999.00066.x.
Artiss T., Schultz T.R., Polhemus D.A., Simon C. Molecular phylogenetic analysis of the dragonfly genera Libellula, Ladona, and Plathemis (Odonata: Libellulidae) based on mitochondrial cytochrome oxidase I and 16S rRNA sequence data (англ.) // Molecular Phylogenetics and Evolution : journal. — Academic Press, 2001. — Vol. 18, no. 3. — P. 348—361. — doi:10.1006/mpev.2000.0867. — PMID11277629.