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Iglésias, S.P., G. Lecointre and D.Y. Sellos. "Extensive paraphylies within sharks of the order Carcharhiniformes inferred from nuclear and mitochondrial genes". = Molecular Phylogenetics and Evolution. — 2005. — Вып. 34. — С. 569–583. — doi:10.1016/j.ympev.2004.10.022.. — PMID15683930.