Cai Chenyang, Tihelka Erik, Giacomelli Mattia, Lawrence John F., Ślipiński Adam, Kundrata Robin, Yamamoto Shûhei, Thayer Margaret K., Newton Alfred F., Leschen Richard A. B., Gimmel Matthew L., Lü Liang, Engel Michael S., Bouchard Patrice, Huang Diying, Pisani Davide and Donoghue Philip C. J. Integrated phylogenomics and fossil data illuminate the evolution of beetles (англ.) // Royal Society open science : Журнал. — London, 2022. — Vol. 9, no. 3. — P. 1—19 (+Suppl.:1—87). — ISSN2054-5703. — doi:10.1098/rsos.211771.
Lubkin S. H., Engel M. S. 2005. Permocoleus, new genus, the first Permian beetle (Coleoptera) from North America. Annals of the Entomological Society of America98: 73—76. doi:10.1603/0013-8746(2005)098[0073:PNGTFP]2.0.CO;2.
Beckemeyer R. J., Engel M. S. 2008. A second specimen of Permocoleus (Coleoptera) from the Lower Permian Wellington formation of Noble County, Oklahoma. Journal of the Kansas Entomological Society81: 4—7. doi:10.2317/JKES-708.01.1.
Cai Chenyang, Tihelka Erik, Giacomelli Mattia, Lawrence John F., Ślipiński Adam, Kundrata Robin, Yamamoto Shûhei, Thayer Margaret K., Newton Alfred F., Leschen Richard A. B., Gimmel Matthew L., Lü Liang, Engel Michael S., Bouchard Patrice, Huang Diying, Pisani Davide and Donoghue Philip C. J. Integrated phylogenomics and fossil data illuminate the evolution of beetles (англ.) // Royal Society open science : Журнал. — London, 2022. — Vol. 9, no. 3. — P. 1—19 (+Suppl.:1—87). — ISSN2054-5703. — doi:10.1098/rsos.211771.
