Uphyrkina, O.; Miquelle, D.; Quigley, H.; Driscoll, C.; O'Brien, S. J. (2002). "Conservation Genetics of the Far Eastern Leopard (Panthera pardus orientalis)". Journal of Heredity. 93 (5): 303–311. doi:10.1093/jhered/93.5.303. PMID12547918.
Uphyrkina, O.; Miquelle, D.; Quigley, H.; Driscoll, C.; O'Brien, S. J. (2002). "Conservation Genetics of the Far Eastern Leopard (Panthera pardus orientalis)". Journal of Heredity. 93 (5): 303–311. doi:10.1093/jhered/93.5.303. PMID12547918.