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Joseph R. Mendelson III; Jay M. Savage; Edgardo Griffith; Heidi Ross; Brian Kubicki; Ronald Gagliardo (2008). "Spectacular new gliding species of Ecnomiohyla (Anura: Hylidae) from Central Panama". Journal of Herpetology. 42 (4): 750–759. doi:10.1670/08-025R1.1. S2CID20233879.