Wanntorp, Livia; Kocyan, Alexander; Renner, Susanne S. (2006). „Wax plants disentangled: A phylogeny of Hoya (Marsdenieae, Apocynaceae) inferred from nuclear and chloroplast DNA sequences”. Molecular Phylogenetics and Evolution. 39 (3): 722—733. PMID16515867. doi:10.1016/j.ympev.2006.01.022.
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Wanntorp, Livia; Kocyan, Alexander; Renner, Susanne S. (2006). „Wax plants disentangled: A phylogeny of Hoya (Marsdenieae, Apocynaceae) inferred from nuclear and chloroplast DNA sequences”. Molecular Phylogenetics and Evolution. 39 (3): 722—733. PMID16515867. doi:10.1016/j.ympev.2006.01.022.