Almeida, Francisca C; Giannini, Norberto P; Simmons, Nancy B; Helgen, Kristofer M (2014). „Each flying fox on its own branch: A phylogenetic tree for Pteropus and related genera (Chiroptera: Pteropodidae)”. Molecular Phylogenetics and Evolution. 77: 83—95. PMID24662680. doi:10.1016/j.ympev.2014.03.009.
Almeida, Francisca C; Giannini, Norberto P; Simmons, Nancy B; Helgen, Kristofer M (2014). „Each flying fox on its own branch: A phylogenetic tree for Pteropus and related genera (Chiroptera: Pteropodidae)”. Molecular Phylogenetics and Evolution. 77: 83—95. PMID24662680. doi:10.1016/j.ympev.2014.03.009.