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André Nel; Patrick Roques; Patricia Nel; Alexander A. Prokin; Thierry Bourgoin; Jakub Prokop; Jacek Szwedo; Dany Azar; Laure Desutter-Grandcolas; Torsten Wappler; Romain Garrouste; David Coty; Diying Huang; Michael S. Engel; Alexander G. Kirejtshuk (2013). „The earliest known holometabolous insects”. Nature. 503 (7475): 257—261. Bibcode:2013Natur.503..257N. PMID24132233. doi:10.1038/nature12629.
André Nel; Patrick Roques; Patricia Nel; Alexander A. Prokin; Thierry Bourgoin; Jakub Prokop; Jacek Szwedo; Dany Azar; Laure Desutter-Grandcolas; Torsten Wappler; Romain Garrouste; David Coty; Diying Huang; Michael S. Engel; Alexander G. Kirejtshuk (2013). „The earliest known holometabolous insects”. Nature. 503 (7475): 257—261. Bibcode:2013Natur.503..257N. PMID24132233. doi:10.1038/nature12629.