K. D. Hill, M. W. Chase, D. W. Stevenson, H. G. Hills, B. Schutzman: The Families and Genera of Cycads: A Molecular Phylogenetic Analysis of Cycadophyta Based on Nuclear and Plastid DNA Sequences. International Journal of Plant Sciences, Band 164, 2003, S. 933–948. doi:10.1086/378538