Almeida, Francisca C; Giannini, Norberto P; Simmons, Nancy B; Helgen, Kristofer M (2014). "Each flying fox on its own branch: A phylogenetic tree for Pteropus and related genera (Chiroptera: Pteropodidae)". Molecular Phylogenetics and Evolution77: 83–95. doi:10.1016/j.ympev.2014.03.009. பப்மெட்:24662680.
Almeida, Francisca C; Giannini, Norberto P; Simmons, Nancy B; Helgen, Kristofer M (2014). "Each flying fox on its own branch: A phylogenetic tree for Pteropus and related genera (Chiroptera: Pteropodidae)". Molecular Phylogenetics and Evolution77: 83–95. doi:10.1016/j.ympev.2014.03.009. பப்மெட்:24662680.